And another thought occurred to me after talking with a friend of mine today. Back in 2014 when the rod let go, I disassembled the engine and found the ding in the combustion chamber mentioned a couple posts back. I assumed that this was due to the piston flying up and hitting the chamber after the rod let go.
But what if the ding in the chamber was due to the piston repeatedly striking the head at high RPM? If I had piston dome to chamber interference for some reason on that cylinder, that would explain why the first rod failed in 2014, when I had been running it successfully in the engine since 2008. In 2014 I put on a new set of heads, caused the piston to chamber interference because of some issue with either the piston or the chamber, and broke the rod. Then, put it all back together again for last year, and the same thing started to happen.
What's more, the ding is on the front side of the chamber, so a fore-aft crack in the rod makes sense under this scenario. This is actually the first really rational explanation for the problem, I think. I can test it by installing last year's head on the short block when I get the short block together, and clay the piston top to check for clearance. Should be enlightening...