I suppose its possible that the skirt was the initial failure, but the scuffing only shows up on three pistons, and all 8 of them show evidence of the support rail moving into the lower ring land; minimum was about 1/4" in, and max was 1/3 of the way around the piston.
Last night I took the one piston I have left here (the rest are now at the machine shop), and re-assembled the oil ring support rail and oil rings after I had filed down the burrs in the lower ring land and also cleaned up the area by the pin so that the dimple in the support rail is pushing against a vertical wall instead of a thin, tapered spot. I pushed on the dimple in the support rail with a screwdriver, trying to get it to move into the ring land, and it wouldn't budge. This leaves me hopeful that this little fix will solve the problem, but I think I'm going to have to re-assemble the engine, run it, and then tear it down again before I know for sure...